‘Science denialism’ is also about outdoor cats

Loss, S. R., Will, T., Longcore, T., & Marra, P. P. (2018). Responding to misinformation and criticisms regarding United States cat predation estimates. Biological Invasions, 20(12), 3385-3396.
Misinformation (or denialism), the disingenuous assertion of information contradicting overwhelming scientific consensus, increasingly poses a challenge for invasion biology. The issue of free-ranging domestic cats (Felis catus) provides an example of this misinformation: overwhelming consensus shows that cats are invasive species that impact wildlife and human health yet free-ranging cat advocates propagate misinformation about such impacts to support policies keeping cats on the landscape. These advocates also attempt to discredit peer-reviewed scientific research on cat impacts, as exemplified by the response to a high-profile paper estimating cats annually kill billions of U.S. birds and mammals (Loss et al. in Nat Commun 4:1396, 2013). Although favorably received by scientific and invasive species management communities, an effort was launched to discredit this paper by criticizing its methods, including a report commissioned by a feral cat advocacy group and a post by a feral cat blogger. These same organizations and individuals have made similar criticisms at scientific conferences and policy roundtables. Given the realized effects of this campaign in influencing invasive species policy, we here respond to these criticisms and show they are characterized by numerous errors and misrepresentations. We conclude that the criticisms are part of the broader campaign to fabricate doubt about outdoor cat impacts and stymie policies favoring removal of cats from the landscape. Because misinformation surrounding cats is emblematic of the broader issue of misinformation and denialism, this response will not only facilitate evidence-based policy for managing cats but also stimulate research and discussion into causes and impacts of misinformation in invasion biology.

Polarized opinions approach on Feral Cat Management

Lorden, R. (2016). Polarized Opinions and Shared Goals: Feral Cat Management in an Academic Community in Kentucky. In Companion Animals in Everyday Life (pp. 183-200). Palgrave Macmillan US.

Although there is general agreement among government agencies, conservation groups and animal welfare organizations that cat populations need to be managed, the management of cats is frequently a topic of debate between biologists/environmental groups and animal welfare/animal rights advocacy groups. The two groups’ beliefs are polarized in regard to impacts on wildlife, management strategies and efficacy of trap-and-neuter (TNR) programs. Concerns of conservation biologists relate to cats as predators, but welfare proponents feel that humans must take responsibility for free-ranging house pets and their feral offspring. It is not surprising that discussions of feral cat management become volatile. Much of the debate hinges upon whether management solutions should use lethal or non-lethal control strategies. Conservation biologists largely support cat euthanasia, while animal welfare activists support non-lethal treatments such as TNR for free-ranging cats. The issues of this larger conservancy/welfare debate framed the discourse of the cat problem and guided solution strategies at Eastern Kentucky University, where a population of cats has existed on the campus for many years. A tentative agreement has been forged between an administration that subscribed to a trap and remove policy, and a network of TNR volunteers. Both sides have focused on the common goal of a reduction in the number of campus cats. In addition, both sides have realized that each setting has its own particular constraints and that solutions must be tailored to those situations. Instead of resorting to invective, data are being collected. When well-meaning people meet and focus on solutions to a shared goal, positive outcomes become possible.

Inefficient TNR

Kilgour, R. J., Magle, S. B., Slater, M., Christian, A., Weiss, E., & DiTullio, M. (2016). Estimating free-roaming cat populations and the effects of one year Trap-Neuter-Return management effort in a highly urban area. Urban Ecosystems, 1-10.

Free-roaming cat populations are increasing in urban areas around the world. Management strategies remain controversial, as attempts to rapidly minimize the impact of cats may conflict with finding an ethical means of population reduction in this domesticated species. Trap-Neuter-Return (TNR) is a non-lethal strategy which can theoretically lead to population decline with an ethical approach. The present study aimed to estimate free-roaming cat populations and also to measure the efficacy of a one-year TNR campaign in a highly urban area. Using a sight/resight methodology, we examined free-roaming cat populations in four sites across two neighbourhoods (Harlem and Bedford-Stuyvesant) in New York City. Sampling was repeated after 1 year, during which an intensive TNR effort occurred in each of those areas. Results from this study found population estimates range from 2.6-4.1 cats/km. Additionally, we found between 78-98% turnover in each study area. After one year of TNR, the proportion of sterilised individuals in our treatment sites increased to 50%. Model results suggest there is no evidence that sterilised individuals are more likely to be encountered, indicating that sterilisation may not affect movement patterns of cats. Free-roaming cats occur at high densities at all study sites, though populations varied, even within the same neighbourhood. We found evidence of considerable migration within study sites, which further complicates the application of a sterilisation management strategy. Management strategies directed toward free-roaming cats, such as TNR, may require a broad-scale approach, involving different facets of the community and should occur over multiple years.

US public opinion on humane treatment of stray cats

Chu, K., & Anderseon, W. M. US Public Opinion on Humane Treatment of Stray Cats. 2007. Alley Cat Allies: Bethesda, MD, 6.

An overwhelming majority of Americans believe that leaving a stray cat outside to live out his life is more humane than having him caught and put down, according to a nationally representative survey conducted for Alley Cat Allies by Harris Interactive in April and May 2007. These results reveal a significant disparity between the public’s humane ethic and the operating policy of most U.S. animal pounds and shelters.
 The current animal control policy is that it is more humane to kill a stray cat now than let him live out his life outdoors; however, this policy rests on untenable bases. While those bases will be discussed in detail below, what is most salient to note is that the socalled humane ethic of the animal control and sheltering system ends more cats’ lives than does any other documented cause of death.

An international comparison of the attitudes of urban people regarding predation by pet cats

Hall, C. M., Adams, N. A., Bradley, J. S., Bryant, K. A., Davis, A. A., Dickman, C. R., ... & Pollock, K. H. (2016). Community Attitudes and Practices of Urban Residents Regarding Predation by Pet Cats on Wildlife: An International Comparison. PLoS One, 11(4), e0151962.

International differences in practices and attitudes regarding pet cats' interactions with wildlife were assessed by surveying citizens from at least two cities in Australia, New Zealand, the UK, the USA, China and Japan. Predictions tested were: (i) cat owners would agree less than non-cat owners that cats might threaten wildlife, (ii) cat owners value wildlife less than non-cat owners, (iii) cat owners are less accepting of cat legislation/restrictions than non-owners, and (iv) respondents from regions with high endemic biodiversity (Australia, New Zealand, China and the USA state of Hawaii) would be most concerned about pet cats threatening wildlife. Everywhere non-owners were more likely than owners to agree that pet cats killing wildlife were a problem in cities, towns and rural areas.

Cat husbandry practices in different countries. (a) Percentage of households that own one, two or more than two cats (b) Percentage of cats kept in different conditions of confinement (c) Percentage of desexed cats. (d) Percentage of cats that have ever caught vertebrate prey.

Agreement amongst non-owners was highest in Australia (95%) and New Zealand (78%) and lowest in the UK (38%). Irrespective of ownership, over 85% of respondents from all countries except China (65%) valued wildlife in cities, towns and rural areas. Non-owners advocated cat legislation more strongly than owners except in Japan. Australian non-owners were the most supportive (88%), followed by Chinese non-owners (80%) and Japanese owners (79.5%). The UK was least supportive (non-owners 43%, owners 25%). Many Australian (62%), New Zealand (51%) and Chinese owners (42%) agreed that pet cats killing wildlife in cities, towns and rural areas was a problem, while Hawaiian owners were similar to the mainland USA (20%).

Percentage agreement of cat owners (dark blue) and non-owners (light blue) in each country to eight survey items: (a) There is a need for cat legislation (b) All cats should be kept in at night time (c) Cats should be kept on their owner's property at all times (d) It is important to have wildlife in cities, towns and rural areas (e) Pet cats killing wildlife in cities, towns and rural areas is a serious problem (f) Pet cats on farms are harmful to wildlife (g) Pet cats in nature reserves are harmful to wildlife (h) Except for a cat owned by a breeder, all cats should be desexed.

Thus high endemic biodiversity might contribute to attitudes in some, but not all, countries. Husbandry practices varied internationally, with predation highest where fewer cats were confined. Although the risk of wildlife population declines caused by pet cats justifies precautionary action, campaigns based on wildlife protection are unlikely to succeed outside Australia or New Zealand. Restrictions on roaming protect wildlife and benefit cat welfare, so welfare is a better rationale.

Use of camera traps to examine the mesopredator release hypothesis in a fragmented landscape

Cove, M. V., Jones, B. M., Bossert, A. J., Clever Jr, D. R., Dunwoody, R. K., White, B. C., & Jackson, V. L. (2012). Use of camera traps to examine the mesopredator release hypothesis in a fragmented Midwestern landscape. The American Midland Naturalist, 168(2), 456-465.

The mesopredator release hypothesis (MRH) has been suggested as a reason why many mammalian generalist mesopredators flourish and become abundant. However, the MRH has only been examined in a limited number of field studies. Some studies have argued that coyotes (Canis latrans) act as top predators in fragmented forest systems and coyote presence has a positive effect on song bird diversity and abundance by controlling mesopredator abundance. We integrated camera trap data and occupancy modeling to determine the factors that affect coyote detection probability and habitat use in a fragmented suburban landscape in central Missouri. We then examined the influence of coyote presence and other habitat variables on mesopredator detection probability and habitat use in the same system. Coyote detection was negatively related to increasing forest cover, whereas red fox (Vulpes vulpes) detection was positively related to increasing urbanization. Coyote occurrence models suggested little habitat selection, while the mesopredator occurrence models suggested an affinity for urbanization. Although there was a slight negative effect of coyote presence on site use by other mesopredators, we suggest that the smaller species are better adapted to coexisting with humans and thus have increased in abundance.

Distribution and correlates of feral cat trapping permits in Los Angeles

Kingsley, G. (2015). Distribution and correlates of feral cat trapping permits in Los Angeles, California (doctoral dissertation, University of Southern California)

Uncontrolled populations of feral cats in urban settings have become of concern to public officials, wildlife scientists, animal rights advocates and the public in general due to the risks they pose to public health, urban wildlife, and esthetics. Solutions to the problem of unmanaged cat populations in cities have been limited in scope by the lack of actual data on feral cats and the urban geographic ranges they occupy. Full extent censuses and environmental analyses have not been collected or performed due to the resources allocations and costs involved. A method for collecting this data without the use of field crews and research summaries exists in the form of unused paper records. Past studies on the problem have used data mining of available records to model cat territories and densities (Aguilar and Farnworth 2012). This approach mitigates the cost while providing information regarding the distributions of these animals. This thesis investigates the spatial properties of feral cat populations in a large metropolitan area (Los Angeles, California) using a previously non-spatialized dataset as a proxy for concentrations of feral cats. The following case study explores two matters: 1) development of a workflow to create a spatial model of feral cat extents from geographic data brought into an analyzable format and 2) analysis of the model data to determine what, if any, variables are correlated with these distributions. The data used for the model were obtained from the City in the form of paper records and successfully imported into a Geographic Information System. Densities of applications were determined from the cleaned and geocoded records and concentrations of both raw density and patterns of clustering were mapped. Modeling of correlations found positive associations with population density and a weak negative correlation with median income. The analysis was assessed and future work on this type of data was considered.

Understanding public perceptions of risk regarding outdoor pet cats to inform conservation action

Gramza, A., Teel, T., VandeWoude, S., & Crooks, K. (2016). Understanding public perceptions of risk regarding outdoor pet cats to inform conservation action. Conservation Biology, 30 (2): 276–286

Free-ranging domestic cats (Felis catus) incur and impose risks on ecosystems and represent a complex issue of critical importance to biodiversity conservation and cat and human health globally. Prior social science research on this topic is limited and has emphasized feral cats even though owned cats often comprise a large proportion of the outdoor cat population, particularly in urban areas. To address this gap, we examined public risk perceptions and attitudes toward outdoor pet cats across varying levels of urbanization, including along the wildland-urban interface, in Colorado, USA. An analysis of 1397 completed surveys showed that residents did not view all types of risks uniformly; they viewed risks of cat predation on wildlife and carnivore predation on cats as more likely than risks of disease transmission to and from wildlife. Additionally, risk perceptions were related to attitudes, prior experiences with cats and cat-wildlife interactions, and cat owner behavior. Findings suggest that changes in risk perceptions may result in behavior change, and they offer insight for communication aimed at promoting risk aversive behaviors and cat management strategies that are acceptable to the public and that directly advance the conservation of native species.

Pathogen exposure varies widely among sympatric populations of wild and domestic felids

Carver, S., Bevins, S. N., Lappin, M. R., Boydston, E. E., Lyren, L. M., Alldredge, M. W., Logan K.A., Sweanor, L.L., Riley, S.P.D., Klein Serieys, L.E., Fisher, R.N., Vickers,T.W., Boyce, W.M., McBride, R., Cunningham, M.C., Jennings, M., Lewis, J.S., Lunn, T. Crooks, K.R. & VandeWoude, S. (2015). Pathogen exposure varies widely among sympatric populations of wild and domestic felids across the United States. Ecological Applications.
Understanding how landscape, host, and pathogen traits contribute to disease exposure requires systematic evaluations of pathogens within and among host species and geographic regions. The relative importance of these attributes is critical for management of wildlife and mitigating domestic animal and human disease, particularly given rapid ecological changes, such as urbanization. We screened >1,000 samples from sympatric populations of puma (Puma concolor), bobcat (Lynx rufus) and domestic cat (Felis catus) across urban gradients in six sites, representing three regions, in North America for exposure to a representative suite of bacterial, protozoal and viral pathogens (Bartonella sp., Toxoplasma gondii, feline herpesvirus-1, feline panleukopenea virus, feline calicivirus, feline immunodeficiency virus). We evaluated prevalence within each species, and examined host trait and land cover determinants of exposure-providing an unprecedented analysis of factors relating to potential for infections in domesticated and wild felids. Prevalence differed among host species (highest for puma and lowest for domestic cat) and was greater for indirectly transmitted pathogens. Sex was inconsistently predictive of exposure to directly transmitted pathogens only, and age infrequently predictive of both direct and indirectly transmitted pathogens. Determinants of pathogen exposure were widely divergent between the wild felid species. For puma, suburban landuse predicted increased exposure to Bartonella sp. in southern California, and FHV-1 exposure increased near urban edges in Florida. This may suggest inter-specific transmission with domestic cats via flea vectors (California) and direct contact (Florida) around urban boundaries. Bobcats captured near urban areas had increased exposure to T. gondii in Florida, suggesting an urban source of prey. Bobcats captured near urban areas in Colorado and Florida had higher FIV exposure, possibly suggesting increased intra-specific interactions through pile-up of home ranges. Beyond these regional and pathogen specific relationships, proximity to the wildland urban interface did not generally increase the probability of disease exposure in wild or domestic felids, emphasizing the importance of local ecological determinants. Indeed, pathogen exposure was often negatively associated with the wildland urban interface for all felids. Our analyses suggest cross-species pathogen transmission events around this interface may be infrequent, but followed by self-sustaining propagation within the new host species.

Ecology and predation of free ranging farm cats

 Kitts-Morgan SE, Caires KC, Bohannon LA, Parsons EI, Hilburn KA (2015) Free-Ranging Farm Cats: Home Range Size and Predation on a Livestock Unit In Northwest Georgia. PLoS ONE 10(4): e0120513. doi:10.1371/journal.pone.0120513

This study’s objective was to determine seasonal and diurnal vs. nocturnal home range size, as well as predation for free-ranging farm cats at a livestock unit in Northwest Georgia. Seven adult cats were tracked with attached GPS units for up to two weeks for one spring and two summer seasons from May 2010 through August 2011. Three and five cats were tracked for up to two weeks during the fall and winter seasons, respectively. Feline scat was collected during this entire period. Cats were fed a commercial cat food daily. There was no seasonal effect (P > 0.05) on overall (95% KDE and 90% KDE) or core home range size (50% KDE). Male cats tended (P = 0.08) to have larger diurnal and nocturnal core home ranges (1.09 ha) compared to female cats (0.64 ha). Reproductively intact cats (n = 2) had larger (P < 0.0001) diurnal and nocturnal home ranges as compared to altered cats. Feline scat processing separated scat into prey parts, and of the 210 feline scats collected during the study, 75.24% contained hair. Of these 158 scat samples, 86 contained non-cat hair and 72 contained only cat hair. Other prey components included fragments of bone in 21.43% of scat and teeth in 12.86% of scat. Teeth were used to identify mammalian prey hunted by these cats, of which the Hispid cotton rat (Sigmodon hispidus) was the primary rodent. Other targeted mammals were Peromyscus sp., Sylvilagus sp. and Microtus sp. Invertebrates and birds were less important as prey, but all mammalian prey identified in this study consisted of native animals. While the free-ranging farm cats in this study did not adjust their home range seasonally, sex and reproductive status did increase diurnal and nocturnal home range size. Ultimately, larger home ranges of free-ranging cats could negatively impact native wildlife.

Cat's impact in Idaho

Hansel, T. E. (2015). Update: Pressing Issues--Impacts of Roaming, Stray, and Feral Domestic Cats (Felis catus) on Birds in Idaho.

Outdoor domestic cats (Felis catus) threaten the welfare of birds and endanger the integrity of the ecosystems into which domestic cats are introduced in Idaho.

Stray Dogs in Houston

Mobility of deers and their arassment by dogs

Progulske, D. R., & Baskett, T. S. (1958). Mobility of Missouri deer and their arassment by dogs. The Journal of Wildlife Management, 184-192.

There is little information on the mobility of the white-tailed deer (Odocoileus virginianus) in the middle United States; data are equally limited on the mobility of white-tails on land that is primarily agricultural or where ther's a yearlong harassment by dogs...

Resolving the urban nest predator paradox: the role of alternative foods for nest predators

Stracey, C. M. (2011). Resolving the urban nest predator paradox: the role of alternative foods for nest predators. Biological Conservation, 144(5), 1545-1552.

Urbanization is a leading cause of species endangerment in the United States; however, certain species thrive in urban habitats. The loss of key predators or the addition of new predators in urban areas could alter the structure of urban communities. A reduction in nest predation is hypothesized to explain the high density of urban birds, yet urban areas typically have increased populations of avian nest predators. The loss of important nest predators in urban habitats, prey switching of urban predators, or successful nest defense against avian nest predators could explain this urban nest predator paradox. To assess these hypotheses I compared nest predation rates of Northern mockingbirds (Mimus polyglottos) in parking lots and residential neighborhoods to populations in pastures and wildlife preserves during 2007–2009 in Florida, USA and placed video cameras on a subset of nests in 2008–2009. Data do not support the hypothesis that urban nest predation rates are consistently lower than non-urban nest predation rates. Of the 56 nest predation events recorded, cats were the dominant urban predator and Cooper’s hawks (Accipiter cooperii) were the dominant non-urban predator. There was no evidence for a loss of important nest predators in urban habitats; however, prey switching by Cooper’s hawks likely occurred. There was also indirect evidence for the importance of nest defense. Furthermore, some of the cats recorded as nest predators in residential neighborhoods were owned cats and all but one cat predation event occurred at night. To reduce nest predation rates, cat owners should keep their cats indoors at night.

Seattle households have more cats than children

Balk, G. 2013. In Seattle, it’s cats, dogs and kids – in that order. Seattle Times. February 1, 2013.

Seattle has more dogs than children. We’ve practically become famous for it. OK, not as famous as we are for coffee or rain, but still. The nation has certainly taken notice. Our canine-to-kid ratio has been written about in USA Today , The New York Times – even the National Postof Canada.It’s not just out-of-towners. Seattleites find it fascinating that we have more dogs than kids, too. The fact has reverberated through the local media since at least 1997, when The Seattle Times first noted that the scales had tipped in favor of dogs here. And it shows no signs of abating; just last year, public radio station KPLU devoted a segment to the phenomenon. Seems like we never tire of talking about it...

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Feeding cats deplates native fauna

Hawkins, C.C., W.E. Grant & M.T. Longnecker. 1999. Effect of subsidized house cats on California birds and rodents. Transactions of the Western Section of The Wildlife Society, 35: 29-33

Cat advocates are establishing feeding stations in public parks, often claiming that well-fed cats pose little threat to wildlife. This claim was tested east of San Francisco, California, in a cat area and a no-cat area. In 1995, more harvest mice were trapped in the no-cat area. In 1996, more harvest mice (Reithrodontomys megalotis) and deer mice (Peromyscus sp.) were trapped in the no-cat area, and more house mice were trapped in the cat area. The numbers of trapped California meadow voles (Microtus californicus) were not different between the areas in either year. More native rodents were trapped in the no-cat area. Birds present during the breeding season were seen more often in the no-cat area. California quail (Callipepla californicus) and California thrashers (Toxostoma redivivum) were present in the no-cat area and absent in the cat area. Cats at artificially high densities, sustained by supplementary feeding, reduced the abundance or native rodent and bird populations, changed the rodent species composition, and may have facilitated the expansion of the house mouse into new areas. Thus we recommend that the feeding of cats in parks should be strictly prohibited. 

Feral cats datasheet

Fitzwater, W.D. 1994. House cats (feral). In S.E. Hygnstrom, R.M. Timm, and G.E. Larson, eds. Prevention and Control of Wildlife Damage. Lincoln: Univ. Neb. Coop. Ext.USDA_APHIS-Wildlife Services Animal Damage Control, and Great Plains Agricultural Council. Pp: C45-C50.

The cat has been the most resistant to change of all the animals that humans have domesticated. All members of the cat family, wild or domesticated, have a broad, stubby skull, similar facial characteristics, lithe, stealthy movements, retractable claws (except the cheetah), and nocturnal habits. Feral cats are house cats living in the wild. They are small in stature, weighing from 3 to 8 pounds, standing 8 to 12 inches high at the shoulder, and 14 to 24 inches long. The tail adds another 8 to 12 inches to their length. Colors range from black to white to orange, and an amazing variety of combinations in between. Cats are found in commensal relationships wherever people are found. In some urban and suburban areas, cat populations equal human populations. In many suburban and eastern rural areas, feral house cats are the most abundant predators. Feral cats prefer areas in and around human habitation. They use abandoned buildings, barns, haystacks, post piles, junked cars, brush piles, weedy areas, culverts, and other places that provide cover and protection. Feral cats are opportunistic predators and scavengers that feed on rodents, rabbits, shrews, moles, birds, insects, reptiles, amphibians, fish, carrion, garbage, vegetation, and leftover pet food. Feral cats feed extensively on songbirds, game birds, mice and other rodents, rabbits, and other wildlife. In doing so, they lower the carrying capacity of an area for native predators such as foxes, raccoons, coyotes, bobcats, weasels, and other animals that compete for the same food base.

Which kills more birds, cats or turbines?

Tim McDonnell on www.motherjones.com

Chart by Tim McDonnell

See also this

Sex bias on cat predation on Eastern bluebirds

Pitts, T. D., & Snow, R. W. (1996). Mortality of Banded Adult Eastern Bluebirds. Audubon Society of Omaha.

Domestic cats are, narrowly, the second most important cause of non-natural mortality, just after shooting. 

Of 51 Eastern Bluebirds killed by domestic cats, 19 were females and 5 were males; the sex of the other birds was not indicated. While this sample size is not much larger than that of bluebirds found dead in the nest (24 versus 16), the trend seen here shows a significant difference in the number of males and females killed (X² = 8.16, df = 1, P <0.005). Figure 5 shows that mortality due to domestic cats is highest during the nesting season. Considering that the female bluebirds, who construct each nest, lay eggs, and incubate the eggs, receive significant amounts of help from the male only when caring for nestlings, we are not surprised that more females are captured by domestic cats. 

(Domestic cats kill an unknown, but probably large, number of nestling bluebirds. However, since this paper deals only with adult mortality we have not explored the effects of cat predation on nestlings.)

Urban cat ownership in Baltimore

Childs, J. E. (1990). Urban cats: their demography, population density, and owner characteristics in Baltimore, Maryland. Anthrozoos: A Multidisciplinary Journal of The Interactions of People & Animals, 3(4), 234-244.

A door-to-door questionnaire survey of 430 households in two areas of Baltimore showed significant differences in cat ownership that were attributable to racial composition. Overall, 29.9% of white households owned a mean of 1.9 cats per cat-owning household, compared to 6.8% of black households owning a mean of 1.0 cat per cat-owning household. The mean age of cats was 3.9 yrs and 1.8 yrs in white and black households, respectively. In contrast, dog ownership in the two areas was nearly identical, with 25.6% of white and 29.1% of black households owning a mean of 1.5 and 1.3 dogs per dog-owning household, respectively. Overall, 62.8% of owned cats were spayed/neutered, 51.8% had received a vaccination in the last year, and 34.7% had been wormed during their lifetimes. A total of 31.6% of households allowed their cats to free-range; 42.5% of the 87 owned cats were in this category. The estimated densities of owned free-ranging cats were 7.0 and 2.8 cats per hectare, for the two areas. Driving counts of free-ranging animals in the two areas suggested a larger population of stray or feral cats in the location with lower reported cat ownership.